A cockatoo is any of the 21 bird species
belonging to the family Cacatuidae. Along with
the Psittacidae family (the true parrots), they
make up the order Psittaciformes. The name
cockatoo originated from the Malay name for
these birds, kaka(k)tua (either from kaka
"parrot" + tuwah, or "older sister" from kakak
"sister" + tua, "old").
Cockatoos share many features with other parrots
including the characteristic curved beak shape
and a zygodactyl foot, with two forward toes and
two backwards toes. They differ, however in a
number of characteristics, including the often
spectacular movable headcrest, the presence of a
gall bladder and some other anatomical details,
and their lack of the Dyck texture feather
composition which causes the bright blues and
greens seen in true parrots. Cockatoo species
are also, on average, larger than the true
parrots (however, the cockatiel is a small
cockatoo and the very large parrots include the
Hyacinth Macaw by length and the Kakapo by
weight). Placement of the cockatoos as a
separate family is fairly undisputed, but it is
not resolved whether or not other living
lineages of parrots (such as the lories) are as
distinct as they are.
Cockatoos have a much more restricted range than
the true parrots, occurring naturally only in
Australia and nearby islands. Eleven of the 21
species exist in the wild only in Australia,
while seven species occur in Indonesia, New
Guinea, and other south Pacific islands. Three
species occur in both New Guinea and Australia.
Contents
1 Cockatoos as endangered or vulnerable species
2 Systematics and evolution
3 Trivia
Cockatoos as endangered or vulnerable species
All species of cockatoo are protected by the
Convention on International Trade in Endangered
Species of Wild Fauna and Flora (also known as
CITES), which makes the import, export and trade
in all wild-caught parrots and cockatoos
illegal.
The following cockatoo species are protected on
the CITES appendix 1 list of endangered species.
* Goffin's cockatoo, Cacatua goffini
* Red-vented Cockatoo, Cacatua haematuropygia
* Moluccan Cockatoo, Cacatua moluccensis
* Yellow-crested Cockatoo, Cacatua sulphurea
o includes the subspecies, Citron-crested
Cockatoo, Cacatua sulphurea citrinocristata
* Palm Cockatoo, Probosciger aterrimus
All of the other cockatoo species are protected
on the CITES appendix 2 list of vulnerable
species.
Systematics and evolution
Phylogeny of the family Cacatuidae based on
Brown & Toft (1999)
Brown & Toft (1999) reviewed the existing
evidence and additional mitochondrial 12S rRNA
sequence data to arrive at a well-supported
phylogeny of the cockatoos. They could
distinguish 3 subfamilies:
1. The all-black Palm Cockatoo represents
distinct lineage that diverged early; it was
previously sometimes grouped with the other
black species but this is incorrect.
2. The dark cockatoos; sexually dichromatic
species which have ample melanin in their
plumage and some red, yellow or orange on wing,
tail and face, barred feathers on wing, tail
and/or body as well as contrasting ear area
spotting in females, while males have the
corresponding feathers unbarred and may lack the
ear spotting. This group includes the remaining
black cockatoos, the Gang-gang Cockatoo and,
interestingly, the cockatiel which had
previously been placed in a subfamily of its own
(Nymphicinae) or even as a broad-tailed parrot.
3. The remaining species, which are all
hypomelanistic and not sexually dimorphic.
The genera Calyptorhynchus and Cacatua can be
further resolved into two subgenera each, and in
the latter case as a distinct third lineage the
white-and-pink Major Mitchell's Cockatoo, which
is intermediate in coloration between the
grey-and-pink Galah and the white Cacatua. It is
best recognized as a monotypic genus Lophocroa.
Indeed, pending further research, all subgenera
could conceivably be raised to species rank.
Intron 7 of nuclear β-fibrinogen sequence data
suggests that the Microglossinae may have
diverged later, and that the cockatiel might be
distinct enough to warrant recognition of the
Nymphicinae (Astuti, 2004), but in other aspects
agrees with the rRNA and newly-interpreted
morphological data.
The fossil record of cockatoos is even more
limited than that of parrots in general, with
only one truly ancient cockatoo fossil known: A
species of Cacatua, most probably subgenus
Licmetis, was found in Early Miocene (16-23 mya)
deposits of Riversleigh, Australia (Boles,
1993). In Melanesia, subfossil bones of Cacatua
species which apparently did not survive early
human settlement were found on New Caledonia and
New Ireland[citation needed]. The bearing of
these fossils on cockatoo evolution and
phylogeny is fairly limited, except that the
Riversleigh fossil allows some tentative dating
of the divergence of subfamilies.
A sulphur-crested cockatoo
A sulphur-crested cockatoo
FAMILY CACATUIDAE
* Subfamily Microglossinae
o Genus Probosciger
+ Palm Cockatoo, Probosciger aterrimus
* Subfamily Calyptorhynchinae - dark cockatoos
o Genus Callocephalon
+ Gang-gang Cockatoo, Callocephalon
fimbriatum
o Genus Nymphicus (tentatively placed here)
+ Cockatiel, Nymphicus hollandicus
o Genus Calyptorhynchus
+ Subgenus Calyptorhynchus - black-and-red
cockatoos
# Red-tailed
Black Cockatoo, Calyptorhynchus (Calyptorhynchus)
banksii
# Glossy
Black Cockatoo, Calyptorhynchus (Calyptorhynchus)
lathami
+ Subgenus Zanda - black-and-yellow/white
cockatoos
#
Yellow-tailed Black Cockatoo, Calyptorhynchus (Zanda)
funereus
#
Short-billed Black Cockatoo, Calyptorhynchus (Zanda)
latirostris
# Long-billed
Black Cockatoo, Calyptorhynchus (Zanda) baudinii
* Subfamily Cacatuinae - white cockatoos
o Genus Eolophus
+ Galah, Eolophus roseicapilla
o Genus Lophocroa
+ Major Mitchell's Cockatoo, Lophocroa
leadbeateri
o Genus Cacatua
+ Subgenus Licmetis - corellas
# Long-billed
Corella, Cacatua (Licmetis) tenuirostris
# Western Corella,
Cacatua (Licmetis) pastinator
# Little Corella,
Cacatua (Licmetis) sanguinea
# Red-vented
Cockatoo, Cacatua (Licmetis) haematuropygia
# Goffin's
Cockatoo, Cacatua (Licmetis) goffini
# Ducorps'
Cockatoo, Cacatua (Licmetis) ducorpsii
+ Subgenus Cacatua - true white
cockatoos
# Sulphur-crested
Cockatoo, Cacatua (Cacatua) galerita
# Yellow-crested
Cockatoo, Cacatua (Cacatua) sulphurea
* Citron-crested Cockatoo, Cacatua (Cacatua)
sulphurea citrinocristata
# Blue-eyed
Cockatoo, Cacatua (Cacatua) ophthalmica
# Moluccan Cockatoo
or Salmon-crested Cockatoo, Cacatua (Cacatua)
moluccensis
# Umbrella
Cockatoo, Cacatua (Cacatua) alba
[edit] Trivia
* A cockatoo is mentioned in John Williamson's
song "True Blue".
* The song, "Like Cockatoos" (in which a sample
of the cockatoos calls can be heard) can be
found on The Cure's 1987 album Kiss Me, Kiss Me,
Kiss Me.
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